Lo, Karin, Thomas, and Queen were all patriarchs.
When they were young and didn't know how to be good patriarchs, the tribes were close to each other, closer than other tribes. Naturally, they often gathered to complain and give each other advice. In winter, whichever tribe had more food would share with the others.
But their stable life was shattered when the Golden Lion tribe wanted to unite with other beastmen to establish a beastman nation.
Because they gullibly believed the lies of the Golden Lion tribe and were betrayed by the Black Wolf tribe, almost all the warriors who participated in that battle sacrificed themselves. Only a few lucky ones survived, but due to severe injuries, their lifespans were greatly shortened, and they passed away one after another not long after.
Lo also sacrificed herself in that battle.
